错误定位:while his unconvincing trade-ship argument overlooks land-based caravans, the overland migration of infected rodents, and the many other animals that carry plague.
正确定位:Twigg employs urban case studies suggesting that the rat population in Europe was both too sparse and insufficiently migratory to have spread plague.
by+过去时间要用过去完成时,这道题为什么没有用过完?