are serving the colony by tending juveniles, gathering food, building the nest, or they battle

are serving the colony in that they tend juveniles, gather food, build the nest, or battle

serve the colony, tending juveniles, gathering food, building the nest, or by battling

serve the colony by tending juveniles, gathering food, by building the nest, or by battling

serve the colony by tending juveniles, gathering food, building the nest, or battling