Ancient hunter-gatherers developed instincts that stigmatized selfishness, and voluntary cooperation was encouraged as well, whether it be within the group or also with outsiders.
selfishness, and voluntary cooperation was encouraged as well, whether it be within the group or
selfishness, encouraging voluntary cooperation as well, both within the group but
selfishness and also encouraging voluntary cooperation, within the group and
selfishness and they encouraged voluntary cooperation, whether it be within the group or
selfishness and encouraged voluntary cooperation, not only within the group but
D: encourage后面跟状语从句,但是whether..or..表示‘或者,或者’的意思。而此处应该是‘不但,而且’的并列关系。
E: 正确。not only..but also做encourage的状语。
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