The army cutworm moth is a critical source of fat for as many as a third of Yellowstone National Park's grizzly bears; they overturn rocks to find them, as many as 40,000 per bear in a single day.
bears; they overturn rocks to find them, as many as
bears; overturning rocks to find the insects, up to
bears, overturning rocks to find them, as many as
bears, and they overturn rocks to find them, up to
bears, which overturn rocks to find the insects, as many as
"who" is used for humans.
sometimes "who" is used for animals that are personified (such as people's pets), but the standard is "which".
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