Before scientists learned how to make a synthetic growth hormone, removing it painstakingly in small amounts from the pituitary glands of human cadavers.
scientists learned how to make a synthetic growth hormone, removing it painstakingly
scientists had learned about making a synthetic growth hormone, they had to remove it painstakingly
scientists learned how to synthesize the growth hormone, it had to be painstakingly removed
learning how to make a synthetic growth hormone, scientists had to remove it painstakingly
learning how to synthesize the growth hormone, it had to be painstakingly removed by scientists
在CD之间选,实际上就是时态的区别
"learning how to synthesize the growth hormone, scientists had to remove it painstakingly" -- dont know if its wrong, but I would still prefer C.
because, of the word "had" which suggests two actions in the past..so ideally, it should be "learned".
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