Birds known as honeyguides exhibit a unique pattern of behavior: the bird leads another animal, such as a honey-badger or a human, to a bees' nest with their chattering when they fly ahead; after the larger animal takes honey, the bird eats the wax and bee larvae.
with their chattering when they fly
with chattering and its flying
by chattering as it flies
by chattering and its flying
by chattering as they are flying
时间连词as。本句在逻辑意思上说的是鸟类在飞行的时候鸣叫,而不是“飞行并且叫”,
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