Studying the fruit fly, a household nuisance but a time-honored experimental subject, has enabled the secrets of how embryos develop to begin to be unraveled by scientists.
Studying the fruit fly, a household nuisance but a time-honored experimental subject, has enabled the secrets of how embryos develop to begin to be unraveled by scientists.
By the study of the fruit fly, a household nuisance and also a time-honored experimental subject, it was possible for the secrets of how embryos develop to begin to be unraveled by scientists.
By studying a household nuisance but a time-honored experimental subject, the fruit fly enabled scientists to begin to unravel the secrets of how embryos develop.
By studying the fruit fly, a household nuisance and also a time-honored experimental subject, the secrets of how embryos develop are beginning to be unraveled by scientists.
The study of the fruit fly, a household nuisance but a time-honored experimental subject, has enabled scientists to begin to unravel the secrets of how embryos develop.
the scientists did the studying., so, the modifier about studying should be as close as possible to "scientists".
("by the study of..." is problematic idiomatically as well.)
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