Scientists believe that unlike the males of most species of moth, the male whistling moths of Nambung, Australia, call female moths to them by the use of acoustical signals, but not olfactory ones, and they attract their mates during the day, rather than at night.
by the use of acoustical signals, but not olfactory ones, and they attract
by the use of acoustical signals instead of using olfactory ones, and attracting
by using acoustical signals, not using olfactory ones, and by attracting
using acoustical signals, rather than olfactory ones, and attract
using acoustical signals, but not olfactory ones, and attracting
RON的解释了 by using 和 using either is fine
by doing sth,blblabla( correct)
passive voice N1 be Ved by N2 (correct)
but by N in choice A of this question is not correct
using 在句尾可以不加逗号直接使用,可以用来当by using 使用