When a prey insect is moved directly toward a beetle that has been chasing it, the beetle immediately turns and runs away without its usual intermittent stopping.

In pursuing a moving insect, the beetles usually respond immediately to changes in the insect's direction, and pause equally frequently whether the chase is up or down an incline.

The beetles maintain a fixed time interval between pauses, although when an insect that had been stationary begins to flee, the beetle increases its speed after its next pause.

If, when a beetle pauses, it has not gained on the insect it is pursuing, the beetle generally ends its pursuit.

When an obstacle is suddenly introduced just in front of running beetles, the beetles sometimes stop immediately, but they never respond by running around the barrier.