Many population studies have linked a high-salt diet to high rates of hypertension and shown that in societies where they consume little salt, their blood pressure typically does not rise with age.
shown that in societies where they consume little salt, their
shown that in societies that have consumed little salt, their
shown that in societies where little salt is consumed,
they showed that in societies where little salt is consumed,
they showed that in societies where they consume little salt, their
in (e), the first "they" is again fine - same antecedent as (d), albeit burdened with the same awkward repetition problem - but the second "they" doesn't have an antecedent.
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