Scientists believe that unlike the males of most species of moth, the male whistling moths of Nambung, Australia, call female moths to them by the use of acoustical signals, but not olfactory ones, and they attract their mates during the day, rather than at night.
by the use of acoustical signals, but not olfactory ones, and they attract
by the use of acoustical signals instead of using olfactory ones, and attracting
by using acoustical signals, not using olfactory ones, and by attracting
using acoustical signals, rather than olfactory ones, and attract
using acoustical signals, but not olfactory ones, and attracting
Either "using" or "by using" would be fine.
Note that "using" is more flexible than other __ing words.
In fact, it's best to think of "using" as belonging to two different categories of words:
1/ It's an __ing word (= it can be used like other __ing words);
2/ It's a preposition, with a meaning largely similar to that of "with".
This is usage #2.
By the way, this distinction is immaterial here, because choice C (= the only choice with "by using") contains blatant non-parallelism.
Takeaways:
— Parallelism is the #1 error in SC. DO NOT think about other things before thinking about parallelism!
— If you're thinking about a small, nit-picky issue, STOP and ask yourself, "Am I being distracted from a larger issue?"
Here, "using"/"by using" is a distraction meant to take your eyes off of the parallelism. Looks like they got you.
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